The game begins with Vito Scaletta (Rick Pasqualone) looking through a photo album, as he begins to tell his story on voiceover. Vito was born in Sicily in 1925 to an extremely poor family. A few years later, he and his family immigrate to the fictional city of Empire Bay (based primarily on New York City but also Chicago) in America, but they are no better off there than they were in Sicily. While in school, Vito gets involved with a local criminal named Joe Barbaro (Robert Costanzo), and they soon become best friends. Vito is arrested during a botched robbery and given a choice: go to jail or join the Army. He chooses the latter and is sent to Sicily during World War II, to assist as a paratrooper in Operation Husky, before being shot and sent home on leave.
Once home, Vito is discharged, courtesy of Joe's mob connections, and learns that his late father Antonio left his mother Maria (Joan Copeland) and sister Francesca (Jeannie Elias) in massive debt. Hoping to make money, Vito turns to Joe, who introduces him to Henry Tomasino (Sonny Marinelli), an inducted member of the Mafia (otherwise known as a made man), working for a mob boss, Alberto Clemente (Nolan North). Working under Henry, Vito is introduced to a real life of crime. Though he is able to pay his father's debt, Vito is soon arrested for illegally distributing ration stamps and sentenced to ten years in prison. While there, he falls in with a crowd led by Leo
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